Variation in eyespot size and colour composition in Bicyclus. The different phenotypes produced by allelic variation in our study locus resemble intra- and interspecific variation in the genus (compare to Figure 1a). The wing pattern of B. anynana lines artificially selected for large eyespots  resembles that of BE individuals. The B. taenias eyespots with broad golden rings resemble those of Fr butterflies. Further work is necessary to determine whether the same alleles or even loci underlie these different phenotypes.