Phylogenetic history and craniofacial morphospace of the Perissodini. (A) Ancestral state reconstruction for foraging habits and diet of the Perissodini, modified from Takahashi et al.. Ancestral foraging preferences are labelled on the tree and preferred habitat depths are indicated to the right. Species are colour coded according to the scatter plots in B and C. Relative warp (RW) analysis revealed several shifts in craniofacial anatomy during Perissodini diversification that correlate with shifts in ecological foraging niches (B, C). (B) Among the Perissodini, RW1 accounts for 39.87% of the variation in shape among species and represents variation in skull length. RW2 accounts for 24.22% of shape variation among species and reflects differences among species in the angle of the mouth. (C) When considering only those species in the genus Perissodus, RW1 accounts for 50.88% of the variation in shape among species and represents variation in skull length. RW2 reflects variation in eye size and accounts for 17.00% of the shape variation.